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This work was supported by funding received by iBB-Institute for Bioengineering and Biosciences (UID/BIO/04565/2013) and other funding (Contracts PTDC/AAC-AMB/111317/2009, Pest-OE/ EQB/LA0023/2011) from European Funds for Economic and Regional Development - FEDERthrough the Competitiveness Factors Operational Programme - COMPETE and National Funds through the Portuguese Foundation of Science and Technology (FCT). M Moreira-Santos thanks to FCT for a postdoctoral fellowship (reference SFRH/BPD/ 99800/2014), funded by Programa Operacional Potencial Humano of QREN Portugal (2014–2020) and by the Portuguese budget through the Ministry of Education and Science.